| Flavour sensors (5,4) |
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| Sample Mr Holly's sensors (9) |
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| When you eat sprouts, what tells you they're palatable (5,4) |
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| Small organs on the tongue (9) |
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| They detect umami etc (5,4) |
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| Detectors of bitterness |
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| They let you sample a dust at its best (5,4) |
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| Sample flowers on the tip of your tongue (5,4) |
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| Umami detectors |
USA Today |
08 Mar 2023 |
| They perform sensationally in oral assessments |
The Telegraph Toughie |
24 Jun 2021 |
| They're on the tip of your tongue |
New York Times |
28 Nov 2020 |
| Dreadful dust beats creating sensational organs |
The Telegraph Toughie |
22 Nov 2018 |
| Umami detectors |
The Washington Post |
16 May 2017 |
| Umami detectors |
LA Times Daily |
16 May 2017 |
| Umami detectors |
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| They are on the tip of your tongue |
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| They may be excited by dinner |
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| They know the difference between sweet and sour |
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| Tongue features (and a hint to this puzzle's theme) |
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| They help distinguish the beginnings of 17-, 32-, 38- and 44-Across |
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| They may be excited by dinner |
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| They help distinguish the beginnings of 17-, 32-, 38- and 44-Across |
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| They're on the tip of your tongue |
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| Sweetness sensors |
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| What caper samplers do? |
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| They may be at the tip of your tongue |
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| These are pleased by 16 Across |
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